NSF 21-506: Division of Integrative Organismal Biology Core Programs
Closed
This grant is closed. A newer notice from the same program is open now: Major Research Instrumentation Program: (MRI) (NSF 27-500). Status: Closes January 29, 2027 (83 days left). It is in our archive of U.S. National Science Foundation notices from 2025.
U.S. National Science Foundation offered grants to any type of organization. It expected to make $70.0 million in total. It is no longer open on Grants.gov.
From the official notice on Grants.gov, last changed June 7, 2025. Checked November 7, 2026.
- Total funding$70.0 million
- Cost sharingNot required
- Funding typeGrant
Other notices with this title
| Notice | Posted | Deadline | Status |
|---|---|---|---|
| NSF 23-547 | June 6, 2025 | Closed | Closed |
| NSF 18-586 | June 6, 2025 | Closed | Closed |
| NSF 20-536 | June 6, 2025 | Closed | Closed |
Key dates
| Posted | June 6, 2025 |
|---|---|
| Archived by Grants.gov | June 7, 2025 |
Who can apply
This opportunity was open to any type of organization.
- Unrestricted (i.e., open to any type of entity above), subject to any clarification in text field entitled "Additional Information on Eligibility"
Is this a realistic fit?
- Most of the program's money (82% over five years) went to colleges and universities.
- No cost sharing or match is required.
What the grant funds
The Division of Integrative Organismal Systems (IOS) Core Programs support research aimed at understanding why organisms are structured the way they are and function as they do. Proposals are welcomed in all of the core scientific program areas supported by the Division of Integrative Organismal Systems (IOS). Areas of inquiry include, but are not limited to, developmental biology and the evolution of developmental processes, nervous system development, structure, modification, function, and evolution; biomechanics and functional morphology, physiological processes, symbioses and microbial interactions, interactions of organisms with biotic and abiotic environments,plant and animal genomics, and animal behavior. Proposals should focus on organisms as a fundamental unit of biological organization. Principal Investigators are encouraged to apply systems approaches that will lead to conceptual and theoretical insights and predictions about emergent organismal properties.
Who has won Biological Sciences grants before
From fiscal 2021 to 2025, Biological Sciences (47.074) awarded $4.2 billion across 6,885 grant awards, an average of $609,810 per award.
| Recipient | State | Awards | Amount |
|---|---|---|---|
| Battelle Memorial Institute | Ohio | 8 | $381 million |
| Arizona State University | Arizona | 57 | $108 million |
| Cornell University | New York | 85 | $85.6 million |
| University of Florida | Florida | 100 | $70.6 million |
| Michigan State University Foundation | Michigan | 105 | $67.9 million |
Applying now
This notice no longer takes applications. Apply under the newer notice, NSF 27-500, which is open now.
The notice on Grants.gov keeps its full text and attachments, which is useful for planning a resubmission.
Questions people ask
Is NSF 21-506 still open?
No. It is no longer open on Grants.gov. A newer notice is open: NSF 27-500, closes January 29, 2027 (83 days left).
Who can apply for Division of Integrative Organismal Biology Core Programs?
U.S. National Science Foundation accepted applications from any type of organization.
What is the deadline for Division of Integrative Organismal Biology Core Programs?
Closed. Applications are no longer accepted; the program's newer notices are on this page when there are any.
How much funding is available?
About $70.0 million in total.
Who has won Biological Sciences grants before?
The largest recipients over five fiscal years were Battelle Memorial Institute, Arizona State University, Cornell University. The program made 6,885 awards worth $4.2 billion in that time.